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+ Introduction
+ History
+ Morphology
+ Terminology
     + Head
     + Antenna
     + Thorax
     + Leg
     + Wing
     + Abdomen
     + Genitalia
     + Chaetotaxy
     + Egg
     + Larva
     + Pupa
     + Addition
     + ESEM
     + Glossary
+ Phylogeny
+ Distribution
+ Ecology
+ Biology
+ Collecting
+ Determination
+ Photography
+ References

Information on Robber Flies
Terminology
Genitalia


Male

Fig. 1: male genitalia, dorsal and lateral
Fig. 1: male genitalia, dorsal and lateral
Dysmachus
  Fig. 1:
1 = male terminalia; postabdomen; hypopygium [1]; male genitalia [6]
2 = abdominal tergite 8, abd tg 8 [1/2]
3 = abdominal sternite 8, abd st 8 [1/2/4]
4 = hypandrium, hypd = abdominal sternite 9 [1/2/3/4/5/6/9]
5 = epandrium (pl. epandria), epand = abdominal tergite 9 [1/2/3/5/6]; sometimes with articulates surstylus (part. in Leptogastrinae) [1/9]; superior forceps [4]
6 = epandrial lobe [3]
7 = cerci (sg. cercus) cerc [1/3/5/9]; proctiger [2/4/6]; surstyli [1]; dorsal: epiproct [1/6]; lamellae [4]; ventral: hypoproct [1/6]; ventral lamella [5] and subanal plate with anus [5]
8 = gonocoxite, goncx [1/2/3/6]; basimere [1]; basistylus [5]; gonocoxa; outer style [#]
9 = gonostylus, gonst [1/3/6/9]; distimere; clasper [1]; dististylus [2/5]; inner style [#]
8+9 = gonopod; anterior gonapophyse; pregonites; harpagone; paramere [1/5]; clasper [4]
10 = aedeagus, aed [1/9]; copulatory organ; intromittent organ; mesosome; penis; phallosome; phallus [1]
Philonicus albiceps, lateral  Philonicus albiceps, ventral
Choerades marginata, lateral  Choerades marginata, posterior

Fig. 2: male genitalia, gonopod
Fig. 2: male genitalia, gonopod
Machimus
  Fig. 2:
11 = gonocoxale apodeme [1/6]
12 = ? [#]
13 = basal lobe [1]
14 = gonocoxale foramen ? [#]
15 = ? [#]
16 = apical lobe [1]; posterior process [#]
17 = ? [#]
18 = ? [#]
19 = ? [#]

Fig. 3: male genitalia, aedeagus
Fig. 3: male genitalia, aedeagus - Apoclea and Grypoctonus
Fig. 3: 20 = aedeagal apodeme, aed apod [1/2/9]; ejaculatory apodeme [6]. - 21 = paramere, pm (also 22) [1/3]; parandrite; paraphyse; penis valves; posterior gonapophyse [1]; paralobe [4]; lateral ejaculatory process [6]; lateral apodeme [9]. - 22 = aedeagal sheath [2/3]; parameral sheath; phallic sheath [6]; paramere sheath [Cannings, 2002]; dorsal and ventral aedeagal sheath (in Leptogastrinae e.g.) [9]. - 23 = basiphallus, bph; phallobase; phallophore; theca [1]. - 24 = distiphallus, distiph; juxta; phallus [1]. - 25 = endoaedeagal process [6]. - 26 = lateral connection column [2]. - 27 = opening of gland [2]. - 28 = endoaedeagus [2]; endoaedeagal tube [Cannings, 2002]. - 29 = gonopore, gonp [1/9]; phallotremata [1]; genital pore [Cannings, 2002]. - 30 = endophallus [1] - non Hennig, 1976; prong [2]; tube [3]. - 31 = ? [#]. - 32 = ventral process [Cannings, 2002]. - 33 = dorsoventral carina [Cannings, 2002]. - 34 = ? [#]. - 35 = ? [#]. - 36 = parameral sheath [6]. - 37 = sperm sac (within the parameral sheath) [6]. - 38 = 22
Euscelidia pallasi, dorsolateral

Female

Euscelidia pallasi, ventral - simple   Choerades marginata, ventral - elongated   Lasiopogon cinctus, dorsolateral - acanthophorites with spines
Philodicus fraterculus, lateral - cerci with spines   Dysmachus trigonus, lateral - sensory hairs
Fig. 4: female genitalia, lateral
Fig. 4: female genitalia, lateral
a. Machimus - b. Dysmachus - c. Laphria - d. Antipalus - e. Stichopogon
  Fig. 4:
  female terminalia; ovipositor [1]
1 = abdominal tergite 7, abd tg 7 [1]
2 = abdominal tergite 8, abd tg 8 [1]
3 = abdominal tergite 9 + 10 ?, abd tg 9+10 [1]
4 = paired cerci (sg. cercus), cerc [1/2]; anal lamellae [2]
5 = abdominal sternite 7, abd st 7 [1]
6 = abdominal sternite 8, abd st 8 [1]; apex in e: Hypogynial valve [Cannings, 2002]
7 = abdominal sternite 9 ?, abd st 9 [1] [abd st 9 is absent in Orthorrhapha sensu Nagatomi & Iwata (1976)]
8 = apex of abdominal tergite 9 [#]
10 = acanthophorite = abdominal tergites 10, which is subdivided into 2 hemitergites [1]; not: abdominal tergite 9 [4]
11 = acanthophorite spines [1/4]

Fig. 5: female genitalia, spermatheca
Fig. 5: female genitalia, spermatheca - Dysmachus [after Theodor, 1980]
  Fig. 5:
  internal structures: spermatheca [1]; receptaculum seminis [8]
1 = reservoir, forms a coiled tube or sclerotized spiral [2]; terminal reservoir [Cannings, 2002]
2 = glands [2]
3 = duct [1/2/8]; reservoir duct [Cannings, 2002]
4 = muscle [2]; muscular sheath [8]
5 = ejection apperatus [2]; spermathecal opening ? [1]
6 = valve of ejection apperatus [2]
7 = apodeme of furca [2]
8 = accessory gland [1/2/8]; appendicular gland; colleterial gland; parovaria [1]
9 = furca (= abd st 9) [2]
10 = common duct [2]
11 = genital chamber [1/8]; vagina [2/8]; bursa copulatrix [8]

References

[1] McAlpine, J.F. (1981): Morphology and terminology - Adults. - In: McAlpine, J.P. et al. (eds.): Manuel of Nearctic Diptera, vol. 1; p. 9-63 - Ottawa: Research Branch, Agriculture Canada, Monograph 27.

[2] Theodor, O. (1980): Fauna Palaestina - Insecta II - Diptera: Asilidae; 446 pp. - Jerusalem: The Israel Academy of Science and Humantities.

[3] Wood, G.C. (1981): Asilidae. - In: McAlpine, J.P. et al. (eds.): Manuel of Nearctic Diptera, vol. 1; p. 549-573 - Ottawa: Research Branch, Agriculture Canada, Monograph 27.

[4] Hull, F.M. (1962): Robber flies of the world, 2 volumes; 907 pp. - Washington: Bulletin of the United States National Museum 224 (1,2).

[5] Artigas, J.N. (1967): The Asilidae (Diptera) of Chile, 2 Vols. - Xerography of Doctoral Diss., University Microfilms: 674 pp.; Ann Arbor.

[6] Sinclair, B.J. & Cumming, J.M. & Wood, D.M. (1994): Homology and phylogenetic implications of male genitalia in Diptera - Lower Brachycera. - Entomologica scandinavica 24: 407-432; Copenhagen.

[7] Reichardt, H. (1929): Untersuchungen über den Genitalapparat der Asiliden. - Zeitschrift für wissenschaftliche Zoologie 135: 257-301; Leipzig.

[8] Snodgrass, R.E. (1935): Principles of insect morphology [reprint 1993]; 667 pp. - Cornell University Press.

[9] Dikow, T. (2001): Revision of the genus Euscelidia Westwood,1850 (Diptera: Asilidae: Leptogastrinae) [part 1] & A phylogenetic analysis of Euscelidia Westwood,1850 (Diptera: Asilidae: Leptogastrinae) based on morphological characters [part 2]. - Rostock: Diploma thesis [unpublished]; 156 pp.

 

 

 

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compiled by: F. Geller-Grimm, contents: F. Geller-Grimm & T. Dikow