Apocleinae
Amblyonychus
Eccritosia
Furcilla
Mallophora
Megaphorus
Nevadasilus
Proctacanthella
Proctacanthus
Promachella
Promachus *
Efferia-group
Albibarbefferia
Aridefferia
Carinefferia
Efferia
Nerax
Pogonioefferia
Triorla
Tuberculefferia

Nearctic species:

1. Promachus albifacies Williston, 1885
2. Promachus aldrichii Hine, 1911
3. Promachus atrox Bromley, 1940
4. Promachus bastardii (Macquart, 1838)
5. Promachus dimidiatus Curran, 1927
6. Promachus fitchii Osten-Sacken, 1878
7. Promachus giganteus Hine, 1911
8. Promachus hinei Bromley, 1931
9. Promachus magnus Bellardi, 1861
10. Promachus minusculus Hine, 1911
11. Promachus nigrialbus Martin in Martin & Papavero, 1970
12. Promachus nigropilosus Schaeffer, 1916
13. Promachus oklahomensis Pritchard, 1935
14. Promachus painteri Bromley, 1934
15. Promachus perpusilla (Walker, 1851)
16. Promachus princeps Williston, 1885
17. Promachus quadratus (Wiedemann, 1821)
18. Promachus rufipes (Fabricius, 1775)
19. Promachus sackeni Hine, 1911
20. Promachus texanus Bromley, 1934
21. Promachus truquii Bellardi, 1861
22. Promachus vertebratus (Say, 1823)

 
 
 
 
 
 
 
 
 

- female -

 
 
 

Unrecognized species: Promachus perpusilla (Walker, 1851) from North America, Promachus magnus Bellardi, 1861 from Mexico (and TX, AZ, NM), and Promachus painteri Bromley, 1934 from Texas (and MEX).
Remarks: A revision is needed for this genus, especially on the base of the internal structures of the genitalia. P. painteri is a true Promachus (Fisher, pers. note, 2005) ###.

	Marginal cell r1 with a gray shadow which sometimes is very narrow.
	Marginal cell r1 without any trace of a gray shadow.

	Male genitalia without silvery hair above. color of the tibiae usually in striking contrast to that of the femora.
	Male genitalia with dense silvery hair above. color of the tibiae very near like that of the femora.

	Gray bands on the dorsum of the abdomen confined to narrow posterior borders of the segments. Gray shadow in the fifth radial cell r5 less than half as wide as the cell. Male genitalia rounded at apex . . . . . Promachus sackeni Hine, 1911 [AZ, UT, NM, TX, MEX]
	Gray bands on the dorsum of the abdomen fully as wide as the black bands.

	Thorax yellowish gray. Male genitalia clothed with gray hair . . . . . Promachus vertebratus (Say, 1823) [MO, MN, IA, NE, CO, WI, MI, OH, PA, IN, IL, KS, OK, SD]
	Thorax deep reddish brown. Male genitalia clothed with black hair.

	Femora reddish or red and black . . . . . Promachus hinei Bromley, 1931 [OH, KS, OH, MO, IL, IN, OK, TX, AR, MS, TN, KY, IA]
	Femora black . . . . . Promachus rufipes (Fabricius, 1775) [IA, IL, IN, OH, PA, MO, MS, KY, TN, AL, GA, SC, NC, WV, VA, FL, DC, AR]

	Abdomen without black hair on any of the segments above.
	Abdomen with black hair on some or all of the first five segments above.

	Abdomen uniformly clothed with short gray hair. Mystax very pale yellow or nearly white. Beard white . . . . . Promachus truquii Bellardi, 1861 [MEX, AZ, NM, TX]
	Abdomen clothed with yellow hair. Mytsax and Beard distinctly yellow.

	Gray shadow in the fifth radial cell r5 wider than the marginal cell r1. Male genitalia almost as long as abdominal segments five, six and seven combined . . . . . Promachus fitchii Osten-Sacken, 1878 [NE, IA, IL, MI, KS, OK, TX, MO, AR, NY, MA, CT, FL, CT, OH, MS, SD, GA, TN, KY, NC]
	Gray shadow in the fifth radial cell r5 distinctly narrower than the marginal cell r1. Male genitalia shorter than abdominal segments five, six and seven combined . . . . . Promachus quadratus (Wiedemann, 1821) [GA, LA, FL], females of Promachus dimidiatus Curran, 1927 [MAN, ALB, SKW, MT, WY, CO, NM, UT, KS, NE, SD, ND, MN, WI, IA], Promachus texanus Bromley, 1934 [TX, OK, MEX], and Promachus oklahomensis Pritchard, 1935 [OK, CO, NE, KS, MO, IL, IN]

	Mystax white or with a very slight shade of yellowish. Legs chiefly black.
	Mystax plainly yellow. legs chiefly yellowish red.

	Abdomen gray pollinose on the sides and extending across narrowly on hind margins of the segments . . . . . Promachus princeps Williston, 1885 [WA, OR, CA, NV, UT]
	Abdomen densely brown pollinose on the sides.

	Wings smoky brown with deep gray shadows in the marginal cell r1 and fifth radial cell r5 . . . . . Promachus atrox Bromley, 1940 [AZ, NM]
	Wings lighter without such deep shadows . . . . . Promachus albifacies Williston, 1885 [AZ, CA, CO, NM, KS, MEX]

	Wings nearly hyaline. Abdomen pale yellowish gray pollinose on the venter and sides . . . . . Promachus quadratus (Wiedemann, 1821) [GA, LA, FL]
	Wings plainly brownish.

	First tergite also with black hairs in males. Abdomen densely brown pollinose on the venter and sides . . . . . Promachus bastardii (Macquart, 1838) [KS, MT, IL, MI, WI, MN, ME, TX, OK, IA, MO, AR, IN, KY, TN, MS, AL, GA, FL, OH, PA, NY, VT, WV, VA, NC, SC, SD]
	First tergite without any black hairs in males.

	Abdomen grayish pollinose on the venter and sides, especially in females (like quadratus) . . . . . Promachus dimidiatus Curran, 1927 [MAN, ALB, SKW, MT, WY, CO, NM, UT, KS, NE, SD, ND, MN, WI, IA]
	Abdomen yellow pollinose on the venter and sides . . . . . Promachus texanus Bromley, 1934 [TX, OK, MEX]

	Legs pure black.
	Legs more or less reddish.

	Abdomen distinctly banded with black and gray, the gray bands slightly more than half as wide as the back . . . . . Promachus nigrialbus Martin in Martin & Papavero, 1970 (= nigripes Hine, 1911) [NM, NV, UT, AZ, TX, MEX]
	Abdomen not distinctly banded, although the narrow posterior margins of the segments are somewhat different from the other parts by appearing whiter . . . . . Promachus aldrichii Hine, 1911 [UT, ID, WA, OR, NV, CA, AZ]

	Very large species, 35-40 mm in length . . . . . Promachus giganteus Hine, 1911 [TX, AZ, NM, MEX]
	Smaller species, not over 25 mm in length.

	Scutum clothed sparsely with short, black hairs. Dorsum of tergites with black hairs . . . . . Promachus nigropilosus Schaeffer, 1916 [CA, OR]
	Scutum clothed with black and white hairs. Dorsum of tergites with black and white hairs . . . . . Promachus minusculus Hine, 1911 [NM, TX, AZ, MEX]

Contents: © Fritz Geller-Grimm, Jorge N. Artigas, 2007
Layout & images: © Fritz Geller-Grimm, 2007